国家转基因生物新品种培育科技重大专项2012ZX08009003-004;国家自然科学基金项目31171537;江苏省高校优势学科建设工程项目PAPD资助。This study was supported by the National Major Project of Breeding for New Transgenic Organisms2012ZX08009003-004;the National Natural Science Foundation of China31171537;the Project Funded by the Priority Academic Program Development of Jiangsu Higher Education Institutions