国家自然科学基金项目31260141;国家林业公益性行业科研专项201104009-08;甘肃省基础研究创新群体项目145RJIG337;甘肃省科技创新服务平台144JTCG254共同资助 Supported by the National Natural Science Foundation of China31260141;National Forestry Public Welfare Profession Scientific Research Project201104009-08;Innovation Group Fund on Fundamental Research of Gansu Province145RJIG337;Science and Technology Innovation Service Platform of Gansu Province144JTCG254